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Nightingale's base skill, Inspiration ( Lady of the Lamp at Elite 2), increases the morale recovery rate of Operators resting in a Dormitory (including Nightingale herself) by 0.1 (0.2 for Lady of the Lamp) when she is assigned to one. Closed Hope can be useful to extend Nightingale's reach and improving her healing efficiency and resistance to Arts damage by a bit.

Nightingale's RIN-X Operator Module, "Closed Hope", extends her healing range by adding a 1-tile extension up front in addition to increasing her ATK and RES once upgraded, White Fiend's Protection also slightly increases the healing received by affected friendlies. While the Mirage is not counted towards the Unit Limit, it loses 3% of its maximum HP every second and Nightingale only have two Mirages available for deployment, which can only be replenished by redeploying her (retreating the Mirages will not replenish their deployment limit), so it must be used wisely. Nightingale's second talent, Fleeting Phantom, allows her to deploy a Mirage on any open melee tiles, which costs 5 DP and has no offensive and defensive capabilities whatsoever, but instead has an unusually high RES of 75 and a 30% chance to dodge Physical attacks, and Taunts enemies (see Aggression for more information), allowing the Mirage to draw enemy fire and potentially protect vulnerable Operators from certain death. Combined with White Fiend's Protection, even Operators without innate RES could have at least 30 RES when Sanctuary is in at least Level 7 and can sometimes dodge incoming Arts attacks, while those with good RES like Angelina or Beeswax can even have at least 100 RES due to the calculation, allowing them to shrug off literally every single Arts attacks (unless if the attack(s) ignore RES, such as Sarkaz Grudgebearers in Windswept Highland with the Objective: Mounting Hatred Contracts)! The third skill, Sanctuary, extends Nightingale's healing range by adding a 1-tile extension up front and 3-tile extensions to the sides, buffs her ATK, and greatly buffs the RES of all friendlies within range (including Nightingale herself) as well as giving them a chance to dodge Arts attacks while active, making it the ultimate anti-Arts skill.The second skill, Arts Shield, has Nightingale generate a short-lasting barrier on the friendlies she heal in the next healing instance which increases their RES and absorbs Arts damage by a portion of Nightingale's ATK, and can stack multiple charges at higher levels.The first skill, Healing Up γ, is an advanced ATK buff which allows Nightingale to heal more HP.Furthermore, all but one of Nightingale's skills allows her to make herself and friendlies withstand Arts damage better than ever: Nightingale's Support specialization is represented by her first talent, White Fiend's Protection, which provides a slight increase of RES to Nightingale herself and friendlies within her range, somewhat mitigating Arts damage they receive and can be useful in the event the player is expected to face enemy Arts attackers. As a Multi-target Medic, Nightingale simultaneously heals up to three friendly units within a range of adjusted 3×3 tiles with a 1-tile extension in front that is extended to adjusted 3×4 tiles at Elite 1 with lower ATK and DP cost when compared to Medical Medics. Nightingale is a 6★ Multi-target Medic who specializes in (obviously) Healing and Support.

A member of the Followers alongside Nearl and Shining, the Sarkaz healer called Nightingale has an obscure past and an emotionless attitude, though it is implied that she was once "a prisoner of her own conscience." Nevertheless, Nightingale, along with the other Followers, joined Rhodes Island to treat other Infected, wherever and whoever they are, with her outstanding knowledge in medical physiology.